杉野 利久Toshihisa Sugino

Last Updated :2024/05/07

所属・職名
大学院統合生命科学研究科　教授
ホームページ
https://home.hiroshima-u.ac.jp/gsbstop/interview/ja/sugino.html
メールアドレス
suginohiroshima-u.ac.jp
その他連絡先
〒739-8528 東広島市鏡山1-4-4 生物生産学部B棟504号室
TEL:082-424-7956 FAX:082-424-7956
自己紹介
酪農業における乳牛飼養管理の問題を栄養，飼育環境からアプローチし，次世代に持続可能な健全性と生産性を両立した乳牛飼養管理技術の開発を目的に，試験研究を行っている。専門領域：家畜飼養学，家畜栄養生理学，乳牛栄養学

基本情報

主な職歴

  • 2021年04月01日, 広島大学, 大学院統合生命科学研究科/生物生産学部, 教授
  • 2015年06月01日, 2019年03月31日, 広島大学, 大学院生物圏科学研究科, 准教授
  • 2007年04月01日, 2015年05月31日, 広島大学, 大学院生物圏科学研究科, 助教
  • 2004年10月04日, 2004年11月03日, 独立行政法人　農業・生物特定産業技術研究機構, 畜産草地研究所, 流動研究員
  • 2003年04月01日, 2007年03月31日, 広島大学, 大学院生物圏科学研究科, 助手

学歴

  • 北里大学, 大学院獣医畜産学研究科, 畜産学専攻, 日本, 2001年04月, 2004年03月
  • 北里大学, 獣医畜産学部, 畜産学科, 日本, 1995年04月, 1999年03月

学位

  • 博士（農学）　(北里大学)
  • 修士（農学）　(北里大学)

教育担当

  • 【学士課程】 生物生産学部 : 生物生産学科 : 応用動植物科学プログラム
  • 【博士課程前期】 スマートソサイエティ実践科学研究院 :
  • 【博士課程前期】 統合生命科学研究科 : 統合生命科学専攻 : 生物資源科学プログラム
  • 【博士課程後期】 スマートソサイエティ実践科学研究院 :
  • 【博士課程後期】 統合生命科学研究科 : 統合生命科学専攻 : 生物資源科学プログラム

担当主専攻プログラム

  • 応用動植物科学プログラム

研究分野

  • 農学 / 動物生命科学 / 動物生産科学

研究キーワード

  • 反芻動物|消化管ホルモン
  • グルカゴン様ペプチド1
  • グレリン
  • 泌乳牛
  • 子牛
  • 代謝

所属学会

  • アメリカ酪農科学会
  • 日本畜産学会, 1999年
  • 国際内分泌学会, 2000年
  • 日本内分泌学会, 2000年
  • 栄養生理研究会, 2001年
  • 関西畜産学会, 2003年

教育活動

授業担当

  1. 2024年, 学部専門, ４ターム, 食料資源論
  2. 2024年, 学部専門, ４ターム, 科学技術倫理学
  3. 2024年, 学部専門, 集中, 基礎物理学実験
  4. 2024年, 学部専門, １ターム, 生理学入門
  5. 2024年, 学部専門, ３ターム, 生物資源科学リサーチフロント
  6. 2024年, 学部専門, ４ターム, 動物栄養学
  7. 2024年, 学部専門, ４ターム, 動物環境生理学
  8. 2024年, 学部専門, 集中, 酪農フィールド科学演習
  9. 2024年, 学部専門, ２ターム, 動物福祉論
  10. 2024年, 学部専門, １ターム, 動物生産生理学
  11. 2024年, 学部専門, ２ターム, 動物生産管理学
  12. 2024年, 学部専門, 集中, 動物生産学実験II
  13. 2024年, 学部専門, 集中, 応用動植物科学特論II
  14. 2024年, 学部専門, 集中, 応用動植物科学特論III
  15. 2024年, 学部専門, セメスター（後期）, 卒業論文I
  16. 2024年, 学部専門, セメスター（前期）, 卒業論文II
  17. 2024年, 学部専門, セメスター（後期）, 卒業論文III
  18. 2024年, 学部専門, 集中, インターンシップI
  19. 2024年, 学部専門, 集中, インターンシップII
  20. 2024年, 学部専門, 集中, 里山フィールド演習
  21. 2024年, 学部専門, 集中, 果樹園芸の里フィールド演習
  22. 2024年, 学部専門, 集中, 牧場実習
  23. 2024年, 学部専門, 集中, 森林フィールド演習
  24. 2024年, 学部専門, 集中, 食品資源フィールド科学演習
  25. 2024年, 学部専門, 集中, 傾斜地フィールド演習
  26. 2024年, 学部専門, 集中, 地域農政特論
  27. 2024年, 学部専門, 通年, 特別実習I
  28. 2024年, 学部専門, 通年, 海洋生物科学技術論と実習IV
  29. 2024年, 学部専門, 通年, 東シナ海学演習
  30. 2024年, 学部専門, 通年, 水産海洋環境学実習I
  31. 2024年, 学部専門, 通年, 水産海洋環境学実習III
  32. 2024年, 学部専門, 通年, 海洋生物科学技術論と実習I
  33. 2024年, 学部専門, 通年, 海洋生物科学技術論と実習II
  34. 2024年, 学部専門, 集中, 水産海洋データ解析演習I
  35. 2024年, 学部専門, 集中, 水産海洋データ解析演習II
  36. 2024年, 学部専門, 集中, 海外生物生産学演習I
  37. 2024年, 学部専門, 集中, 海外生物生産学演習II
  38. 2024年, 学部専門, １ターム, 植物育種学
  39. 2024年, 学部専門, セメスター（後期）, 植物病理学
  40. 2024年, 学部専門, セメスター（前期）, Graduate Thesis I
  41. 2024年, 学部専門, セメスター（後期）, Graduate Thesis II
  42. 2024年, 学部専門, セメスター（前期）, Graduate Thesis III
  43. 2024年, 学部専門, セメスター（後期）, Graduate Thesis IV
  44. 2024年, 学部専門, 集中, 国際課題研究
  45. 2024年, 学部専門, 集中, International Applied Biological Sciences I
  46. 2024年, 学部専門, 集中, (AIMS) Molecular-Level Understanding of Functionality of Foods
  47. 2024年, 学部専門, ３ターム, Molecular-Level Understanding of Functionality of Foods
  48. 2024年, 学部専門, 集中, (AIMS) Introduction Physiology of Domestic Animals
  49. 2024年, 学部専門, ３ターム, Introductory Physiology of Domestic Animals
  50. 2024年, 修士課程・博士課程前期, １ターム, 生物資源科学特別演習Ａ
  51. 2024年, 修士課程・博士課程前期, ２ターム, 生物資源科学特別演習Ａ
  52. 2024年, 修士課程・博士課程前期, ３ターム, 生物資源科学特別演習Ｂ
  53. 2024年, 修士課程・博士課程前期, ４ターム, 生物資源科学特別演習Ｂ
  54. 2024年, 修士課程・博士課程前期, 年度, 生物資源科学特別研究
  55. 2024年, 修士課程・博士課程前期, ３ターム, 応用動物生産論

研究活動

学術論文（★は代表的な論文）

  1. Response of plasma glucagon-like peptide-2 to feeding pattern and intraruminal administration of volatile fatty acids in sheep, DOMESTIC ANIMAL ENDOCRINOLOGY, 60巻, pp. 31-41, 2017
    DOIを用いた論文検索結果へのリンク
  2. Effects of partial replacement of corn grain with lactose in calf starters on ruminal fermentation and growth performance, JOURNAL OF DAIRY SCIENCE, 100巻, 8号, pp. 6177-6186, 2017
    DOIを用いた論文検索結果へのリンク
  3. ★, Plasma concentrations of glucagon-like peptide 1 and 2 in calves fed calf starters containing lactose., Journal of Dairy Science, 2017
  4. GH secretory responses to ghrelin and GHRH in growing and lactating dairy cattle, Domestic Animal Endocrinology, 28巻, pp. 34-45, 20050401
  5. Effects of short-term exposure to heat stress on splanchnic metabolism in sheep, J. Anim. Feed Sci., 13巻, pp. 359-362, 20040401
  6. Ghrelin enhances glucose-induced insulin secretion in scheduled meal-fed sheep, Journal of Endocrinology, 189巻, pp. 67-75, 20060401
  7. 同位体アミノ酸をトレーサーとしたウマの大腿部筋の筋タンパク質合成および分解速度測定方法の確立, 馬の科学, 43巻, 1号, pp. 8-18, 20060401
  8. 運動負荷したサラブレッドにおける同位体アミノ酸（L-[ring-2H5]- phenylalanine）標識法による後肢大腿部筋タンパク質の合成および分解の測定；2コンポーネントメソッド（動脈と静脈濃度差法）, 馬の科学, 43巻, 4号, pp. 267-282, 20060401
  9. Effects of nutritional status on hormone concentrations of the somatotropin axis and metabolites in plasma and colostrum of Japanese Black cows, ANIMAL SCIENCE JOURNAL, 88巻, 4号, pp. 643-652, 2017
    DOIを用いた論文検索結果へのリンク
  10. Ghrelin differentially modulates glucose-induced insulin secretion according to feeding status in sheep, JOURNAL OF ENDOCRINOLOGY, 194巻, 3号, pp. 621-625, 2007
    DOIを用いた論文検索結果へのリンク
  11. Ghrelin differentially modulates the GH secretory response to GHRH between the fed and fasted states in sheep, DOMESTIC ANIMAL ENDOCRINOLOGY, 37巻, 1号, pp. 55-60, 2009
    DOIを用いた論文検索結果へのリンク
  12. Plasma hormone and metabolite concentrations involved in the somatotropic axis of Japanese Black heifers in association with growth hormone gene polymorphism, DOMESTIC ANIMAL ENDOCRINOLOGY, 37巻, 4号, pp. 243-249, 2009
    DOIを用いた論文検索結果へのリンク
  13. Changes in blood pancreatic polypeptide and ghrelin concentrations in response to feeding in sheep, JOURNAL OF ANIMAL SCIENCE, 88巻, 6号, pp. 2103-2107, 2010
    DOIを用いた論文検索結果へのリンク
  14. Effects of glucose and amino acids on ghrelin secretion in sheep, ANIMAL SCIENCE JOURNAL, 81巻, 2号, pp. 199-204, 2010
    DOIを用いた論文検索結果へのリンク
  15. Effects of amino acids infused into the vein on ghrelin-induced GH, insulin and glucagon secretion in lactating cows, ANIMAL SCIENCE JOURNAL, 82巻, 2号, pp. 267-273, 2011
    DOIを用いた論文検索結果へのリンク
  16. Comparisons of insulin related parameters in commercial-type chicks: Evidence for insulin resistance in broiler chicks, PHYSIOLOGY & BEHAVIOR, 103巻, 2号, pp. 233-239, 20110303
    DOIを用いた論文検索結果へのリンク
  17. Follicular Persistence Induced by Adrenocorticotropic Hormone Administration in Goats, JOURNAL OF REPRODUCTION AND DEVELOPMENT, 57巻, 2号, pp. 212-216, 2011
    DOIを用いた論文検索結果へのリンク
  18. Plasma ghrelin concentration is decreased by short chain fatty acids in wethers, DOMESTIC ANIMAL ENDOCRINOLOGY, 41巻, 1号, pp. 50-55, 2011
    DOIを用いた論文検索結果へのリンク
  19. Differential localization of lingual antimicrobial peptide in the digestive tract mucosal epithelium of calves, VETERINARY IMMUNOLOGY AND IMMUNOPATHOLOGY, 142巻, 42737号, pp. 87-94, 20110715
    DOIを用いた論文検索結果へのリンク
  20. Effects of high ambient temperature on urea-nitrogen recycling in lactating dairy cows, ANIMAL SCIENCE JOURNAL, 82巻, 4号, pp. 531-536, 2011
    DOIを用いた論文検索結果へのリンク
  21. Central and peripheral effects of glucagon-like peptide 1 (GLP-1) on food intake and plasma concentrations of glucose and insulin in the goats., S Afr J Anim Sci (ISRP), 29巻, pp. 14-15, 19990401
  22. ★, A transient ghrelin surge occurs just before feeding in a scheduled meal-fed sheep., Biochem. Biophys. Res. Commun., 295巻, pp. 255-260, 20020401
  23. ★, A transient surge of ghrelin secretion before feeding is modified by different feeding regimens in sheep., Biochem. Biophys. Res. Commun., 298巻, pp. 785-788, 20020401
  24. Circulating leptin response to feeding and exogenous infusion of insulin in sheep exposed to thermoneutral and cold environments., Comp. Biochem. Physiol. A., 134巻, pp. 329-335, 20030401
  25. ★, Involvement of cholinergic neurons in the regulation of the ghrelin secretory response to feeding in sheep., Biochem. Biophys. Res. Commun., 304巻, pp. 308-312, 20030401
  26. ★, Effects of ghrelin on food intake and neuroendocrine function in sheep, Animal Reproduction Science, 82-83巻, pp. 183-194, 20040801
  27. 反芻動物のグレリン分泌と作用, 栄養生理研究会報, 48巻, 1号, pp. 19-34, 20040401
  28. Effect of amino acid and glucose adoministration following exercise on the turnover of muscle protein in the hindlimb femoral region of thoroughbreds, Equine Veterinary Journal, 36巻, pp. 611-616, 20060401
  29. Postprandial changes in plasma concentrations of growth hormone and ghrelin around weaning in the goat, General and Comparative Endocrinology, 148巻, pp. 368-374, 20060421
  30. Effect of potato pulp as a dietary carbohydrate source on nitrogen excretion and urea metabolism in sheep, Energy and Protein Metabolism and Nutrition, pp. 435-436, 20070901
  31. Ghrelin differentially modulates glucose-induced insulin secretion according to feeding status in sheep., Journal of Endocrinology, 194巻, pp. 621-625, 20070628
  32. Plasma levels of ghrelin and glucagon-like peptide 1 in periparturient Holstein dairy cows., Proc. 12th AAAP, 20060922
  33. Effect of potato pulp as a dietary carbohydrate source on nitrogen excretion and urea metabolism in sheep., Energy and Protein Metabolism and Nutrition, pp. 435-436, 20070916
  34. Net Portal Fluxes of Nitrogen Metabolites in Holstein Steers Fed Diets Containing Different Dietary Ratios of Whole-crop Corn Silage and Alfalfa Hay, ASIAN-AUSTRALASIAN JOURNAL OF ANIMAL SCIENCES, 22巻, 3号, pp. 371-377, 20090301
  35. Insulin Dynamics in Transition Dairy Cows as Revealed by Intravenous Glucose Tolerance Testing, JOURNAL OF ANIMAL AND VETERINARY ADVANCES, 9巻, 18号, pp. 2333-2337, 20100101
  36. 周産期乳牛における乳腺組織での血漿アミノ酸の動静脈差の推移, 広大FSC報告, 7巻, pp. 19-29, 20100401
  37. Chemical structure of nuclear proteins which are phosphorylated during meiotic maturation of starfish oocytes, BIOCHEMISTRY, 39巻, 21号, pp. 6390-6400, 20000530
    DOIを用いた論文検索結果へのリンク
  38. Effects of ghrelin on food intake and neuroendocrine function in sheep, ANIMAL REPRODUCTION SCIENCE, 30011巻, pp. 183-194, 2004
    DOIを用いた論文検索結果へのリンク
  39. Effects of short-term exposure to heat stress on splanchnic metabolism in sheep, JOURNAL OF ANIMAL AND FEED SCIENCES, 13巻, pp. 359-362, 2004
    DOIを用いた論文検索結果へのリンク
  40. Ghrelin enhances glucose-induced insulin secretion in scheduled meal-fed sheep, JOURNAL OF ENDOCRINOLOGY, 189巻, 1号, pp. 67-75, 2006
    DOIを用いた論文検索結果へのリンク
  41. Postprandial changes in plasma concentrations of growth hormone and ghrelin around weaning in the goat, GENERAL AND COMPARATIVE ENDOCRINOLOGY, 148巻, 3号, pp. 368-374, 20060915
    DOIを用いた論文検索結果へのリンク
  42. Net Portal Fluxes of Nitrogen Metabolites in Holstein Steers Fed Diets Containing Different Dietary Ratios of Whole-crop Corn Silage and Alfalfa Hay, ASIAN-AUSTRALASIAN JOURNAL OF ANIMAL SCIENCES, 22巻, 3号, pp. 371-377, 2009
    DOIを用いた論文検索結果へのリンク
  43. The effect of dietary ratios of corn silage and alfalfa hay on carbohydrate digestion and retention time of feed particles in the gastrointestinal tract of steers, ANIMAL SCIENCE JOURNAL, 80巻, 5号, pp. 546-555, 2009
    DOIを用いた論文検索結果へのリンク
  44. Insulin Dynamics in Transition Dairy Cows as Revealed by Intravenous Glucose Tolerance Testing, JOURNAL OF ANIMAL AND VETERINARY ADVANCES, 9巻, 18号, pp. 2333-2337, 2010
    DOIを用いた論文検索結果へのリンク
  45. Effects of calcium salts of long-chain fatty acids and rumen-protected methionine on plasma concentrations of ghrelin, glucagon-like peptide-1 (7 to 36) amide and pancreatic hormones in lactating cows, DOMESTIC ANIMAL ENDOCRINOLOGY, 42巻, 2号, pp. 74-82, 2012
    DOIを用いた論文検索結果へのリンク
  46. Effects of starch content of calf starter on growth and rumen pH in Holstein calves during the weaning transition, JOURNAL OF DAIRY SCIENCE, 95巻, 8号, pp. 4478-4487, 2012
    DOIを用いた論文検索結果へのリンク
  47. Effects of feeding a calf starter on molecular adaptations in the ruminal epithelium and liver of Holstein dairy calves, JOURNAL OF DAIRY SCIENCE, 95巻, 5号, pp. 2585-2594, 2012
    DOIを用いた論文検索結果へのリンク
  48. Plasma concentrations and effects of glucagon-like peptide-1 (7-36) amide in calves before and after weaning, DOMESTIC ANIMAL ENDOCRINOLOGY, 43巻, 4号, pp. 299-306, 2012
    DOIを用いた論文検索結果へのリンク
  49. Effects of intravenous ghrelin injection on plasma growth hormone, insulin and glucose concentrations in calves at weaning, ANIMAL SCIENCE JOURNAL, 84巻, 4号, pp. 310-315, 2013
    DOIを用いた論文検索結果へのリンク
  50. Effects of glucose and volatile fatty acids on blood ghrelin concentrations in calves before and after weaning, JOURNAL OF ANIMAL SCIENCE, 90巻, 13号, pp. 4839-4845, 2012
    DOIを用いた論文検索結果へのリンク
  51. Effects of forage intake level on nitrogen net flux by portal-drained viscera of mature sheep with abomasal infusion of an amino acid mixture, ANIMAL, 7巻, 10号, pp. 1614-1621, 2013
    DOIを用いた論文検索結果へのリンク
  52. Energy metabolism by splanchnic tissues of mature sheep fed varying levels of lucerne hay cubes, ANIMAL, 7巻, 10号, pp. 1622-1630, 2013
    DOIを用いた論文検索結果へのリンク
  53. Ingestion of medium chain fatty acids by lactating dairy cows increases concentrations of plasma ghrelin, DOMESTIC ANIMAL ENDOCRINOLOGY, 45巻, 4号, pp. 216-223, 2013
    DOIを用いた論文検索結果へのリンク
  54. Effects of dietary combination of corn and rice as whole crop silage and grain sources on carbohydrate digestion and nitrogen use in steers, ANIMAL SCIENCE JOURNAL, 85巻, 2号, pp. 127-134, 2014
    DOIを用いた論文検索結果へのリンク
  55. Effects of calcium salts of medium-chain fatty acids on plasma metabolite and hormone concentrations in early lactating dairy cows, ANIMAL PRODUCTION SCIENCE, 54巻, 10号, pp. 1699-1702, 2014
    DOIを用いた論文検索結果へのリンク
  56. Effect of glucagon-like peptide-1 and ghrelin on liver metabolites in steers, ANIMAL PRODUCTION SCIENCE, 54巻, 10号, pp. 1732-1736, 2014
    DOIを用いた論文検索結果へのリンク
  57. Effects of ethanol on splanchnic nutrient metabolism in sheep at different intake levels.?, Energy and Protein Metabolism and Nutrition in Sustainable Animal Production, pp. 441-442, 201309
  58. 泌乳牛における栄養素と血漿グレリン濃度の関係について., 栄養生理研究会報 57:33-44., 57巻, pp. 33-44, 201305
  59. A preliminarily study for predicting body weight and milk properties in lactating Holstein cows using a three-dimensional camera system, COMPUTERS AND ELECTRONICS IN AGRICULTURE, 111巻, pp. 186-193, 2015
    DOIを用いた論文検索結果へのリンク
  60. Insulin-independent actions of glucagon-like peptide-1 in wethers, ANIMAL SCIENCE JOURNAL, 86巻, 4号, pp. 385-391, 2015
    DOIを用いた論文検索結果へのリンク
  61. Milk production, plasma metabolite profiles and mammary arterial-venous differences of milk precursors in early lactation cows milked at different frequencies by an automatic milking system, ANIMAL SCIENCE JOURNAL, 86巻, 5号, pp. 499-507, 2015
    DOIを用いた論文検索結果へのリンク
  62. Effects of fat-enriched diet and methionine on insulin sensitivity in lactating cows, JOURNAL OF ANIMAL SCIENCE, 93巻, 6号, pp. 2778-2784, 2015
    DOIを用いた論文検索結果へのリンク
  63. Effects of dietary combination of corn and rice as whole crop silage and grain sources on carbohydrate digestion and nitrogen use in steers., ANIMAL SCIENCE JOURNAL, 2014
  64. Effects of calcium salts of medium-chain fatty acids on plasma metabolite and hormone concentrations in early lactating dairy cows., ANIMAL PRODUCTION SCIENCE, 2014
  65. Effect of glucagon-like peptide-1 and ghrelin on liver metabolites in steers., ANIMAL PRODUCTION SCIENCE, 2014
  66. A preliminarily study for predicting body weight and milk properties in lactating Holstein cows using a three-dimensional camera system., Computers and Electronics in Agriculture, 2015
  67. A comparison of plasma glucose and oxidative status in lactating dairy cows in summer and autumn, ANIMAL SCIENCE JOURNAL, 87巻, 10号, pp. 1212-1217, 2016
    DOIを用いた論文検索結果へのリンク
  68. Metabolomic profiling reveals differential effects of glucagon-like peptide-1 and insulin on nutrient partitioning in ovine liver, ANIMAL SCIENCE JOURNAL, 87巻, 12号, pp. 1480-1489, 2016
    DOIを用いた論文検索結果へのリンク
  69. Identification of Ghrelin in Fertilized Eggs of Chicken, JOURNAL OF POULTRY SCIENCE, 46巻, 3号, pp. 257-259, 2009
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  134. Insulin Dynamics in Transition Dairy Cows as Revealed by Intravenous Glucose Tolerance Testing, JOURNAL OF ANIMAL AND VETERINARY ADVANCES, 9巻, 18号, pp. 2333-2337, 2010
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  144. Milk production, plasma metabolite profiles and mammary arterial-venous differences of milk precursors in early lactation cows milked at different frequencies by an automatic milking system, ANIMAL SCIENCE JOURNAL, 86巻, 5号, pp. 499-507, 2015
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  154. Postruminal supply of amino acids enhances ghrelin secretion and lipid metabolism in feed-deprived sheep, ANIMAL SCIENCE JOURNAL, 89巻, 12号, pp. 1663-1672, 2018
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  157. Evaluating the rumen-protected lysine stability in forage-based total mixed rations in vitro and determining the lysine Brix value, ANIMAL SCIENCE JOURNAL, 90巻, 8号, pp. 932-938, 2019
  158. Effect of varying fermentation conditions with ensiling period and inoculum on photosynthetic pigments and phytol content in Italian ryegrass (Lolium multiflorum Lam.) silage, ANIMAL SCIENCE JOURNAL, 2019
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受賞

  1. 2010年03月28日, 日本畜産学会奨励賞, 社団法人日本畜産学会, 「反芻動物のグレリン分泌と作用に関する研究」において、その内容が高く評価され、家畜の栄養生理に関する研究に多大な貢献があったと認定
  2. 2006年09月, XIIth AAAP Animal Science Congress 2006 Excellent Presentation Award, "Prof. Inkee Paik, Ph.D, President", PLASMA LEVELS OF GHRELIN AND GLUCAGON-LIKE PEPTIDE 1 IN PERIPARTURIENT HOLSTEIN DAIRY CATTLE
  3. 2004年03月, 日本畜産学会大会優秀発表賞, 日本畜産学会理事長, ヒツジのグレリン分泌機構、特にインスリンの分泌抑制作用の可能性について
  4. 2000年03月, 日本畜産学会優秀ポスター賞, 粗飼料又は濃厚飼料給与時の血中GLP-1濃度の変化