Last Updated :2024/06/06

Affiliations, Positions
Graduate School of Integrated Sciences for Life, Associate Professor
Other Contact Details
1-4-4 Kagamiyama, Higashi-Hiroshima 739-8528, Japan
TEL : (+81) FAX : (+81)
My research interests include the taxonomy and systematics of algae, in particular seaweed taxonomy of tropical regions such as Southeast Asia.

Basic Information

Major Professional Backgrounds

  • 2008/10/01, 2016/03/31, Hiroshima University, Graduate School of Biosphere Science, Associate Professor (Special Appointment)
  • 2016/04/01, 2019/03/31, Hiroshima University, Graduate School of Biosphere Science, Associate Professor
  • 2019/04/01, Hiroshima University, Graduate School of Integrated Sciences for Life, Associate Professor

Academic Degrees

  • Ph.D. (Biology), University of North Carolina at Chapel Hill
  • Master of Science (Marine Biology), University of the Philippines

Research Keywords

  • algae, freshwater Rhodophyta, museum studies, phycology, phytogeography, seaweeds

Affiliated Academic Societies

  • Association of Systematic Biologists of the Philippines
  • International Phycological Society
  • Japanese Society of Phycology

Educational Activity

Course in Charge

  1. 2024, Liberal Arts Education Program1, 2Term, Human and Ecological Systems in Transition
  2. 2024, Undergraduate Education, 2Term, Introduction to Applied Biological Science I
  3. 2024, Undergraduate Education, Intensive, Laboratory Work in Hydrospheric BiologyIII
  4. 2024, Undergraduate Education, 2Term, Hydrospheric Environmental ScienceII
  5. 2024, Undergraduate Education, Second Semester, Graduation Thesis I
  6. 2024, Undergraduate Education, First Semester, Graduation Thesis II
  7. 2024, Undergraduate Education, Second Semester, Graduation Thesis III
  8. 2024, Undergraduate Education, Intensive, Overseas Exercise of Applied Biological Science I
  9. 2024, Undergraduate Education, Intensive, (AIMS) Global Environmental Issues and Managements
  10. 2024, Undergraduate Education, Intensive, Global Environmental Issues and Managements
  11. 2024, Graduate Education (Master's Program) , 1Term, English Presentation Methods in Science and Technology
  12. 2024, Graduate Education (Master's Program) , 3Term, Sustainable Marine Environment
  13. 2024, Graduate Education (Master's Program) , Second Semester, Interdisciplinary Seminar A
  14. 2024, Graduate Education (Doctoral Program) , Second Semester, Interdisciplinary Seminar B
  15. 2024, Graduate Education (Master's Program) , 3Term, Sustainable Marine Environment

Research Activities

Academic Papers

  1. Marine algae of the Sulu Sea Islands, Philippines II: annotated list of the brown seaweeds (Phaeophyceae) from the Cuyo Islands, PHYTOTAXA, 152(1), 1-17, 20131211
  2. An early 18th century account of marine algae from the Philippines., Philippine Journal of Science, 113-117, 2013
  3. Redefining the role of natural history collections and museums in an era of global climate change., Philippine Journal of Systematic Biology, 53-62, 2013
  4. Moderate thermal stress causes active and immediate expulsion of photosynthetically damaged zooxanthellae (Symbiodinium) from corals, PLOS ONE, 9(12), 20141210
  6. Conspecificity of two morphologically distinct calcified red algae from the northwest Pacific Ocean: Galaxaura pacifica and G. filamentosa (Galaxauraceae, Rhodophyta), BOTANICAL STUDIES, 54, 2013
  7. Phylogeny of Betaphycus (Gigartinales, Rhodophyta) as inferred from COI sequences and morphological observations on B-philippinensis, JOURNAL OF APPLIED PHYCOLOGY, 26(1), 587-595, 2014
  8. Chlorophycean parasite on a marine fish, Sillago japonica (Japanese sillago), PARASITOLOGY INTERNATIONAL, 62(6), 586-589, 2013
  9. Foliose Halymenia species (Halymeniaceae, Rhodophyta) from Southeast Asia, including a new species, Halymenia malaysiana sp nov., BOTANICA MARINA, 58(3), 203-217, 2015
  10. Macrobenthic algae of the southern coasts of Catanduanes Island, Philippines: Historical account and critical review of records., Thailand Natural History Museum Journal, 9(1), 95-108, 2015
  11. Enhancement of Marine Phytoplankton Growth by Steel-making Slag as a Promising Component for the Development of Algal Biofuels, ISIJ INTERNATIONAL, 56(4), 708-713, 2016
  12. Succession of seawall algal communities on artificial substrates, Raffles Bulletin of Zoology, 32(32), 1-10, 20160506
  13. A simple and rapid determination method for zooxanthellal genetic diversity in giant clams using multiplex PCR, FISHERIES SCIENCE, 82(5), 747-753, 2016
  14. Marine algae of the Sulu Sea islands, Philippines III. Taxonomic account of the Gracilariaceae (Rhodophyta) from the Cuyo Islands, PALAWAN SCIENTIST, 10, 1-16, 201807
  15. A genetic diversity assessment of Halymenia malaysiana (Halymeniaceae, Rhodophyta) from Malaysia and the Philippines based on COI-5P and rbcL sequences, JOURNAL OF APPLIED PHYCOLOGY, 30(6), 3445-3454, 201812
  16. First Report of the Red Algal Genus Chondria C. Agardh (Rhodomelaceae, Rhodophyta) for the Marine Flora of Bangladesh, DIVERSITY-BASEL, 11(6), 201906
  17. Caulerpa falcifolia Harvey and Bailey (Chlorophyta) from Sibutu Island, Tawi-Tawi, a new record for the marine algal flora of the Philippines., Tropical Natural History, 19, 1-7, 20190201
  18. New record of Grateloupia taiwanensis SM Lin et HY Liang in Vietnam: Evidence of morphological observation and rbcL sequence analysis., Biodiversitas, 20, 688-695, 20190630
  19. New record of the red alga, Endosiphonia horrida, in Singapore., Singapore Biodiversity Records, 2019(51), 89-90, 20190831
  20. Participation of female farmers groups in kai algal processing and production in northern Thailand., Human and Environmental Security in the Era of Global Risks: Perspectives from Africa, Asia and the Pacific Islands., 265-274, 20190201
  21. Allelopathic effects of macroalgae on Pocillopora acuta coral larvae, MARINE ENVIRONMENTAL RESEARCH, 151, 201910
  22. Satellite image analysis reveals changes in seagrass beds at Van Phong Bay, Vietnam during the last 30 years, AQUATIC LIVING RESOURCES, 33, 20200505
  23. Effects of water temperature, light and nitrate on the growth of sporelings of the non-geniculate coralline alga Lithophyllum okamurae (Corallinales, Rhodophyta), JOURNAL OF APPLIED PHYCOLOGY, 32(3), 1923-1931, 202006
  24. The Ulvophyceae (Chlorophyta) of eastern Sorsogon, Philippines, including Halimeda magnicuneata sp. nov. (Bryopsidales), BOTANICA MARINA, 63(5), 439-453, 202010
  25. Sterol and fatty acid profiles of three macroalgal species collected from the Chabahar coasts, southeastern Iran, AQUACULTURE INTERNATIONAL, 29(1), 155-165, 202102
  26. Visia cylindrocellularis: A new record of freshwater red algae (Rhodophyta) in Singapore and extension of its geographical distribution range, NOVA HEDWIGIA, 112(3-4), 323-333, 202105
  27. Potential bioactive compounds as SARS-COV-2 inhibitors from extracts of the marine alga Halymenia durvillei (Rhodophyta) - a computational study., Arabian Journal of Chemistry, 11(103393), 20210911

Invited Lecture, Oral Presentation, Poster Presentation

  1. Dead men tell no tales but dead plants do: Herbarium specimens provide clues about recent environmental changes, 2018/04/27, With Invitation, English, National University of Singapore, National University of Singapore, Herbarium specimens deposited in museums around the world have traditionally been used to support taxonomic studies. Lately, new uses have been identified on top of their traditional roles. Some histological and chemical features therein have been used to describe and relate with changing environmental conditions, providing proxy data potentially useful for studies in climate and environmental change. In this presentation, examples from flowering plants and marine algae showing variability in phenology and morphological characteristics including their presence or absence have been used for mapping general community changes through time and space, biogeographical shifts, invasion history, urbanization and other anthropogenic impacts, affording a record of the recent past even if sketchy and perhaps, a peek into the future.


  1. 2016/11/05, Excellent Educator Award, Hiroshima University
  2. 2005/08/15, Most Outstanding Alumnus in Marine Science, University of San Carlos Alumni Association

Social Activities

History as Committee Members

  1. International Exchange Committee, 2019/04, 2021/03
  2. Public Relations Committee, 2019/04, 2021/03

History as Peer Reviews of Academic Papers

  1. Phycological Research, Others
  2. Phytotaxa, Others
  3. The Philippine Scientist, Chief editor
  4. Philippine Journal of Science, Others, Reviewer, 3
  5. Philippine Journal of Systematic Biology, Chief editor
  6. Journal of Applied Phycology, Others, reviewer, 2
  7. 2019, Botanica Marina, Others, reviewer, 2