Hiroyuki Horiuchi

Last Updated :2019/07/05

Affiliations, Positions
Graduate School of Integrated Sciences for Life, ., Professor
E-mail
hhori10hiroshima-u.ac.jp

Basic Information

Major Professional Backgrounds

  • 1992/01/01, 2002/03/31, Hiroshima University, Faculty of Applied Biological Science, Research Assistant
  • 2007/04/01, Hiroshima University, Graduate School of Biosphere Science, Research associate
  • 2010/04/01, Hiroshima University, associate professor
  • 2012/05/01, Hiroshima University, professor

Educational Backgrounds

  • Hiroshima University, Graduate School, Division of Biophere Sciences, Japan, 1988/04, 1990/03
  • Hiroshima University, Faculty of Applied Biological Science, Japan, 1983/04, 1988/03

Academic Degrees

  • Doctor of Science, Hiroshima University
  • Master of Agriculture, Hiroshima University

Educational Activity

  • 【Bachelor Degree Program】School of Applied Biological Science : Department of Applied Biological Science
  • 【Master's Program】Graduate School of Integrated Sciences for Life : Division of Integrated Sciences for Life : Program of Food and AgriLife Science
  • 【Doctoral Program】Graduate School of Integrated Sciences for Life : Division of Integrated Sciences for Life : Program of Food and AgriLife Science

In Charge of Primary Major Programs

  • Applied Molecular and Cellular Biology

Research Fields

  • Agricultural sciences;Animal life science;Integrative animal science

Research Keywords

  • chicken
  • thrombocyte
  • Cytokine
  • membrane protein
  • embryonic stem cells
  • pluripotent stem cell
  • germ cell
  • genome editing
  • dementia

Affiliated Academic Societies

  • The Molecular Biology Society of Japan, 1994/04
  • The Japanease Society for Genome Editing, 2016/04

Educational Activity

Course in Charge

  1. 2019, Undergraduate Education, 1Term, Basic Molecular Biology
  2. 2019, Undergraduate Education, Intensive, Laboratory Work in General Biology I
  3. 2019, Undergraduate Education, 2Term, Reading of Foreign Literature
  4. 2019, Undergraduate Education, 3Term, Immunobiology
  5. 2019, Undergraduate Education, Intensive, Laboratory Work in Immunobiology
  6. 2019, Undergraduate Education, 2Term, Animal Cell Technology
  7. 2019, Undergraduate Education, Academic Year, Graduation Thesis
  8. 2019, Undergraduate Education, First Semester, Project-Based Learning for Active Learninng
  9. 2019, Undergraduate Education, First Semester, Exective Seminar (3)
  10. 2019, Undergraduate Education, Intensive, Overseas Training
  11. 2019, Undergraduate Education, Intensive, Laboratory Exercise (1)
  12. 2019, Undergraduate Education, Intensive, Laboratory Exercise (2)
  13. 2019, Graduate Education (Master's Program) , First Semester, Practice in Molecular and Applied Biosciences
  14. 2019, Graduate Education (Master's Program) , Second Semester, Research for Academic Degree Dissertation
  15. 2019, Graduate Education (Master's Program) , 2Term, Applied Immunology
  16. 2019, Graduate Education (Master's Program) , 4Term, Exercises in Applied Immunology
  17. 2019, Graduate Education (Master's Program) , 1Term, Exercises in Food andAgriLife Science A
  18. 2019, Graduate Education (Master's Program) , 2Term, Exercises in Food andAgriLife Science A
  19. 2019, Graduate Education (Master's Program) , 3Term, Exercises in Food andAgriLife Science B
  20. 2019, Graduate Education (Master's Program) , 4Term, Exercises in Food andAgriLife Science B
  21. 2019, Graduate Education (Master's Program) , Academic Year, Research for Academic Degree Dissertation in Food andAgriLife Science
  22. 2019, Graduate Education (Master's Program) , 2Term, Animal Life Science I
  23. 2019, Graduate Education (Master's Program) , 4Term, Animal Life Science II
  24. 2019, Graduate Education (Doctoral Program) , Academic Year, Research for Academic Degree Dissertation in Integrated Life Sciences

Research Activities

Academic Papers

  1. Comparison of sex determination mechanism of germ cells between birds and fish: Cloning and expression analyses of chicken forkhead box L3-like gene., Developmental Dynamics, Open access(Open access), 1-11, 20190617
  2. Zymosan A enhances humoral immune responses to soluble protein in chickens, JOURNAL OF VETERINARY MEDICAL SCIENCE, 79(8), 1335-1341, 201708
  3. Expression of ayu antimicrobial peptide genes after LPS stimulation, JOURNAL OF VETERINARY MEDICAL SCIENCE, 79(6), 1072-1080, 2017
  4. Induction of non-specific suppression in chicks by specific combination of maternal antibody and related antigen, JOURNAL OF VETERINARY MEDICAL SCIENCE, 77(11), 1363-1369, 2015
  5. Studies on Morphology and Cytochemistry in Blood Cells of Ayu Plecoglossus altivelis altivelis, JOURNAL OF VETERINARY MEDICAL SCIENCE, 76(5), 693-704, 2014
  6. Manipulating the avian epiblast and epiblast-derived stem cell, Methods Mol. Biol., 1074, 151-173, 201311
  7. Molecular cloning of chicken interleukin-5 receptor alpha-chain and analysis of its binding specificity, DEVELOPMENTAL AND COMPARATIVE IMMUNOLOGY, 37(42798), 354-362, 2012
  8. Characteristics of Novel Chicken Embryonic Stem Cells Established Using Chicken Leukemia Inhibitory Factor, JOURNAL OF POULTRY SCIENCE, 48(1), 64-72, 2011
  9. Construction of an Insertion Vector for Gene Targeting of Chicken Lens-specific Gene, JOURNAL OF POULTRY SCIENCE, 47(2), 144-148, 2010
  10. Allergenicity study of EGFP-transgenic chicken meat by serological and 2D-DIGE analysis, FOOD AND CHEMICAL TOXICOLOGY, 48(5), 1302-1310, 2010
  11. Induction of immune suppression in the chick by an optimal dose of an immunizing antigen in the presence of its specific maternal antibody., J. Vet. Med. Sci., 72(3), 276-262, 20090601
  12. Prolonged suppression of chick humoral immune response by antigen specific maternal antibody., J. Vet. Med. Sci., 71(4), 417-424, 20090401
  13. Generation and characterization of chicken monoclonal antibodies against human LOX-1., MAbs., 1(4), 357-363, 20090401
  14. Chicken antibody against a restrictive epitope of prion protein distinguishes normal and abnormal prion proteins., Biologicals, 35(4), 303-308, 20071025
  15. Cloning of the chicken interleukin-13 receptor alpha 2 gene and production of a specific monoclonal antibody., Developmental and Comparative Immunology, 31(4), 394-406, 20070510
  16. Development of recombinant chicken IgY from single chain fragment of variable region for diagnosis of BSE., Biologicals, 35(1), 31-34, 20070301
  17. Antibody engineering and transgenic technology in chickens, Food and Food Ingredients Journal of Japan, 211(11), 948-954, 20061101
  18. Effect of novel monoclonal antibodies on LIF-induced signaling in chicken blastodermal cells., Dev. Comp. Immunol., 30(5), 513-522, 20060401
  19. Characterization and expression analysis of a chicken interleukin-6 receptor alpha., Dev. Comp. Immunol., 30(4), 419-429, 20060401
  20. Maintenance of chicken embryonic stem cells in vitro., In Methods in Molecular Biology (Embryonic Stem Cell Protocols; Second Edition Vol. 1), 329, 17-34, 20060401
  21. Humanization of chicken monoclonal antibody using phage-display system., Mol. Immunol., 43(6), 634-642, 20060201
  22. Tributyltin disturbs bovine adrenal steroidogenesis by two modes of action., Steroids, 70(14), 913-921, 20051201
  23. Inhibition of prion propagation in scrapie-infected mouse neuroblastoma cell lines using mouse monoclonal antibodies against prion protein., Biochem. Biophys. Res. Commun., 335(1), 197-204, 20050901
  24. Stable production of recombinant chicken antibody in CHO-K1 cell line., Biologicals, 33(3), 169-174, 20050901
  25. Biological activity of recombinant chicken interleukin-6 in chicken hybridoma cells., Vet. Immunol. Immunopathol., 106(42737), 97-105, 20050601
  26. Generation of a mouse monoclonal antibody against chicken interleukin-6., Hybridoma (Larchmt)., 24(2), 115-117, 20050501
  27. Excision of foreign gene product with cathepsin D in chicken hepatoma cell line., Biochem. Biophys. Res. Commun., 330(2), 533-539, 20050501
  28. Characterization and expression analysis of the chicken interleukin-11 receptor alpha chain., Dev. Comp. Immunol., 29(4), 349-359, 20050401
  29. Genomic organization of the chicken T-cell receptor beta chain D-J-C region., The Journal of Veterinary Medical Science, 66(12), 1509-1515, 20041201
  30. Expression vectors for chicken-human chimeric antibodies, Biologicals, 32(4), 213-218, 20041201
  31. Cloning and characterization of a chicken leukemia inhibitory factor, Avian Immunology Research Group, 8, 80, 20040904
  32. Characteruzation and expression analysis of the chicken interleukin-11 receptor alpha., Avian Immunology Research Group, 8, 82, 20040904
  33. Biological activities of recombinant chicken interleukin-6 on chicken hybridoma cells., Avian Immunology Research Group, 8, 85, 20040904
  34. Establishment of a chicken monoclonal antibody panel against mammalian prion protein, The Journal of Veterinary Medical Science, 66(7), 807-814, 20040801
  35. Chicken leukemia inhibitory factor maintains chicken embryonic stem cells in the undifferentiated state, The Journal of Biological Chemistry, 279(23), 24513-24520, 20040601
  36. Molecular cloning and characterization of chicken tumor necrosis factor (TNF)-superfamily ligands, CD30L and TNF-related apoptosis inducing ligand (TRAIL), The Journal of Veterinary Medical Science, 66(6), 643-650, 20040601
  37. A monoclonal antibody against chicken thrombocytes reacts with the cells of thrombocyte lineage, The Journal of Veterinary Medical Science, 66(3), 243-250, 20040301
  38. Molecular study on chicken tumor necrosis factor receptor-II and tumor necrosis factor receptor-associated factor-5., Vet. Immunol. Immunopathol., 98(42737), 31-41, 20040301
  39. Chicken peripheral blood CD3+CD4-CD8- cells are regulated by endocrine and immune systems., J. Vet. Med. Sci., 66(2), 143-148, 20040201
  40. Immunobiology of chicken germinal center: II. Accumulation of apoptotic cells within the germinal center, Cell Tissue Res., 314(2), 215-221, 20031101
  41. Two expression vectors for the phage-displayed chicken monoclonal antibody, Journal of Immunological Methods, 280(42737), 157-164, 20030901
  42. Flow cytometric analysis of chicken NK activity and its use on the effect of restraint stress, Journal of Veterinary Medical Science, 65(9), 995-1000, 20030901
  43. Generation of antibodies against prionprotein by scrapie-infected cell immunization of PrP0/0 mice, HYBRIDOMA and HYBRIDOMICS, 22(4), 263-266, 20030801
  44. Development of Peyerユs Patch and Cecal Tonsil in Gut-Associated Lymphoid Tissues in the Chicken Embryo., J. Vet. Med. Sci., 65(5), 607-614, 20030501
  45. Immunobiology of chicken germinal center: I. Changes in surface Ig class expression in the chicken splenic germinal center after antigen stimulation, Developpmental and Comparative Immunology, 27(2), 159-166, 20030201
  46. Studies on chicken cytokines: cloning and application, 46(2), 137-149, 20021201
  47. Cloning and characterization of a chicken platelet-derived growth factor B-chain cDNA., Dev. Comp. Immunol., 26(1), 73-83, 20020101
  48. Establishment of monoclonal antibodies against chicken GP130, Veterinary Immunology Symposium, 6, 106, 20010715
  49. Characterization and expression of three forms of cDNA encoding chicken platelet-derived growth factor-A chain, Gene, 272(42737), 181-190, 20010711
  50. Role of bursin in the development of B lymphocytes in chicken embryonic Bursa of Fabricius, Developpmental and Comparative Immunology, 25(42861), 485-493, 20010601
  51. CD3+CD4-CD8- cells present in chicken peripheral blood are regulated by aging and stress, CURRENT PROGRESS ON AVIAN IMMUNOLOGY RESEARCH, 6, 315-321, 20001008
  52. Construction of recombinant chicken monoclonal antibody using phage display system, CURRENT PROGRESS ON AVIAN IMMUNOLOGY RESEARCH, 6, 335-340, 20001001
  53. Construction of recombinant monoclonal antibodies from a chicken hybridoma line secreting specific antibody., Cytotechnology, 32, 191-198, 20000401
  54. cDNA cloning and mRNA expression of PDGF in chicken thrombocyte, Proceedings of the International Veterinary Cytokine and Vaccine Conference, 234-237, 20000316
  55. Expression of chicken gp130 mRNA in various chicken organs and cells, Proceedings of the International Veterinary Cytokine and Vaccine Conference, 330-333, 20000316
  56. Chicken monoclonal antibodies against synthetic bovine prion protein peptide., J. Vet. Med. Sci., 60(6), 777-779, 19980601
  57. Manipulation of chick embryo and production of transgenic chicken., 3(4), 39-44, 19980401
  58. Ontogeny of thrombocytes in chicken embryos, The challenges of Lower Vertebrates to Immunology, 99, 19971113
  59. Two chicken B cell lines resistant to ouabain for the production of chicken monoclonal antibodies, The Journal of Veterinary Medical Science, 58(11), 1053-1056, 19961101
  60. Identification of the gene encoding VP7 of serotype G3 equine rotavirus, Journal of Equine Science, 7(1), 17-20, 19960401
  61. Detection of growth factor(s) in chicken thrombocytes., Avian Immunology Research Group, 19940320
  62. Vitronectin, 201-208, 19930401
  63. Establishment of chicken B hybridomas and their applications, Avian Immunology in Progress, 91-96, 19930401
  64. Identification of fibronectin in chicken thrombocytes, Cell Structure and Function, 17(2), 93-98, 19920417
  65. Vibrio virulence factors and the quantitative analysis of cytotoxiicity elaborated by environmental isolates, Cytobios, 66(266-267), 143-151, 19910401
  66. Preliminary evidence of growth factor(s) from chicken thrombocytes- Growth effects of chicken fibroblasts cultures, Japanese Journal of Veterinary Science, 52(3), 559-565, 19900601

Invited Lecture, Oral Presentation, Poster Presentation

  1. Analyses of germ-cell fate determination in chickens, Hiroaki Matsumoto, Ryo Ezaki, Ryota Kajihara, Mei Matsuzaki, Shuichi Furusawa, Hiroyuki Horiuchi, 4th Annual Meeting of the Japanese Society for Genome Editing, 2019/06/05, Without Invitation, The Japanese Society for Genome Editing, Tokyo
  2. Elucidation of mechanism of chicken sex determination using genome editing, Yuki Okaza, Mayuko Ymawaki, Ryo Ezaki, Mai Matsuzaki, Shuichi Furusawa, Hiroyuki Horiuchi, 4th Annual Meeting of the Japanese Society for Genome Editing, 2019/06/04, Without Invitation, The Japanese Society for Genome Editing, Tokyo
  3. Study on biomarkers for diagnosis of rapidly progressive dementias, Yusuke Sanuki, Yumi Tanaka, Ryo Ezaki, Shuichi Furusawa, Noriyuki Nishida, Katsuya Satoh and Hiroyuki Horiuchi, The 41st Annual Meeting of the Molecular Biology Society of Japan, 2018/11/30, Without Invitation, The Molecular Biology Society of Japan, Yokohama
  4. Forced-expression of target gene by insertion of promoter using PITCh system, Kosuke Yasui, Ryo Ezaki, Shuichi Furusawa and Hiroyuki Horiuchi, The 41st Annual Meeting of the Molecular Biology Society of Japan, 2018/11/29, Without Invitation, The Molecular Biology Society of Japan, Yokohama
  5. Comparison of sex determination mechanism of germ cells between birds and fish by analyses of chicken foxl3., Kennosuke Ichikawa, Ryo Ezaki, Shuichi Furusawa and Hiroyuki Horiuchi, The 41st Annual Meeting of the Molecular Biology Society of Japan, 2018/11/28, Without Invitation, The Molecular Biology Society of Japan, Yokohama
  6. Germ-cell differentiation induction of chicken pluripotent stem cells., Sachi Uehata, Asako Hirano, Ryo Ezaki, Shuichi Furusawa and Hiroyuki Horiuchi, The 41st Annual Meeting of the Molecular Biology Society of Japan, 2018/11/28, Without Invitation, The Molecular Biology Society of Japan, Yokohama
  7. Functional analysis of DAZL in avian primordial germ cells using genome-editing tool, Kennosuke Ichikawa, Haruto Masaki, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 3rd Annual Meeting of the Japanese Society for Genome Editing, 2018/06/20, Without Invitation, The Japanese Society for Genome Editing, Hiroshima city
  8. Function analyses of sex determination related gene in chickens, Chikako Yamada, Yuki Nakagawa, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 3rd Annual Meeting of the Japanese Society for Genome Editing, 2018/06/20, Without Invitation, The Japanese Society for Genome Editing, Hiroshima city
  9. Functional analysis of DAZL in avian primordial germ cells using genome-editing tool, Haruto Masaki, Kennosuke Ichikawa, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 3rd Annual Meeting of the Japanese Society for Genome Editing, 2018/06/19, Without Invitation, The Japanese Society for Genome Editing, Hiroshima city
  10. Application of MMEJ-assisted gene knock-in by PITCh system in chickens, Ryo Ezaki, Fumiko Kameyama, Shuichi Furusawa, Hiroyuki Horiuchi, The 3rd Annual Meeting of the Japanese Society for Genome Editing, 2018/06/19, Without Invitation, The Japanese Society for Genome Editing, Hiroshima city
  11. Functional analysis of DAZL in avian primordial germ cells, Haruto Masaki, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, ConBio2017, 2017/12/08, Without Invitation, The Molecular Biology Society of Japan, Kobe, JAPAN
  12. Study on avian sex determination using genome editing, Yuki Okaza, Ryo Ezaki, shuichi Furusawa, Hiroyuki Horiuchi, ConBio2017, 2017/12/08, Without Invitation, The Molecular Biology Society of Japan, Kobe, JAPAN
  13. Isolation, culture and characterization of chicken amniotic mesenchymal stem cells, Karina Saheki, Ryo Ezaki, Fumiya Hirose, Shuichi Furusawa and Hiroyuki Horiuchi, ConBio2017, 2017/12/07, Without Invitation, The Molecular Biology Society of Japan, Kobe, JAPAN
  14. Elucidation of mechanism of avian sex determination using genome editing, Yuki Okaza, Yuki Nakagawa, Ryo Ezaki, shuichi Furusawa, Hiroyuki Horiuchi, The 2nd Annual Meeting of the Japanese Society for Genome Editing, 2017/09/29, Without Invitation, the Japanese Society for Genome Editing, Osaka
  15. Cloning and expression analyses of chicken forkhead box L3, Kennosuke Ichikawa, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The Fourth World Congress of Reproductive Biology, 2017/09/27, Without Invitation, The Society for Reproduction and Development, Okinawa, Jaoan
  16. Study on biomarkers for diagnosis of prion disease, Yumi Tanaka, Ryo Ezaki, Shuichi Furusawa, Noriyuki Nishida, Haruo Matsuda, Katusya Satoh, Hiroyuki Horiuchi, The 39th Annual Meeting of the Molecular Biology Society of Japan, 2016/12/01, Without Invitation, Molecular Biology Society of Japan, Yokohama, Japan
  17. An improved protocol for stable and simple culturing of chicken primordial germ cells using apoptosis inhibitors, Ryo Ezaki, Fumiya Hirose, Shuichi Furusawa and Hiroyuki Horiuchi, The 39th Annual Meeting of the Molecular Biology Society of Japan, 2016/12/01, Without Invitation, Molecular Biology Society of Japan, Yokohama, Japan
  18. Are chicken epiblast-derived stem cells naïve or primed pluripotent state?, Asako Hirono, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 39th Annual Meeting of the Molecular Biology Society of Japan, 2016/11/30, Without Invitation, Molecular Biology Society of Japan, Yokohama, Japan
  19. Cloning and expression analyses of chicken forkhead domain box L3, Kennosuke Ichikawa, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 39th Annual Meeting of the Molecular Biology Society of Japan, 2016/11/30, Without Invitation, Molecular Biology Society of Japan, Yokohama, Japan
  20. Genome editing in chicken primordial germ cell using genome editing tools, Yuki Nakagawa, Ryo Ezaki, Fumiya Hirose, Shuichi Furusawa, Tetsushi Sakuma, Asato Kuroiwa, Takashi Yamamoto, Hiroyuki Horiuchi, The 1st Annual Meeting of the Genome Editing Society of Japan, 2016/09/06, Without Invitation, The Japanese Society of Genome Editing, Hiroshima, Japan
  21. Genome editing in chicken epiblast derived stem cells using CRISPR/Cas9, Fumiko Kameyama, Ryo Ezaki, Shuichi Furusawa, Hiroyuki Horiuchi, The 1st Annual Meeting of the Genome Editing Society of Jaopan, 2016/09/06, Without Invitation, The Japanese Society of Genome Editing, Hiroshima. Japan
  22. Establishment and preservation of embryonic stem cells from Onagadori, Teruo Maeda, Qian Sun, Ryo Ezaki, Hiroyuki Horiuchi, The 17th Asian-Australasian Association of Animal Production Societies Animal Science Congress, 2016/08/22, Without Invitation, Fukuoka, JAPAN
  23. Stable and simple culture protocol for chicken primordial germ cells using apoptosis inhibitor, Ryo Ezaki, Fumiya Hirose, Shuichi Furusawa, Hiroyuki Horiuchi, The 17th Asian-Australasian Association of Animal Production Societies Animal Science Congress, 2016/08/22, Without Invitation, Fukuoka, JAPAN
  24. Establishment of quail ES-like cells, Mikiharu Nakano, Saki Soeda, Hiroyuki Horiuchi, Yoichi Matsuda, 2016 PSA Annual Meeting, 2016/07/11, Without Invitation, New Orleans, Louisiana, USA
  25. Establishment of high-sensitivity detection method of FABP3 in cerebrospinal fluid of CJD patients, Yumi Tanaka, Ryo Ezaki, Shuich Furusawa, Noriyuki Nishida, Haruo Matsuda, Katusya Satoh, Hiroyuki Horiuchi, PRION 2016, 2016/05/10, Without Invitation, Tokyo, JAPAN
  26. Establishment of quail ES-like cells, Mikiharu Nakano, Saki Soeda, Hiroyuki Horiuchi, Yoichi Matsuda, Avian Model Systems 9, 2016/03/28, Without Invitation, Taipei, TAIWAN
  27. Study on male sex determination gene in the bird using genome editing technology., Yuki Nakagawa, Ryo Ezai, Tetsushi Sakuma, Asato Kuroiwa, Takashi Yamamoto, Hiroyuki Horiuchi., Biochemistry and Molecular Biology 2015, 2015/12/04, Without Invitation, Kobe, JAPAN

Awards

  1. 2011年10月26日, 10th Transgenic Technology (TT) meeting,Best Poster Award, Dr. Lluis Montoliu ISTT Executive Committee, President, Florida (USA), October 24-26 (2011)で開催された10th Transgenic Technology (TT) meetingでポスター発表を行い,発表の内容,ポスターのレイアウト等が高く評価
  2. 2015年12月03日, BMB2015 Young-Scientist Presentation Award, The Molecular Biology Society of Japan, Japanese Biochemical Society, Study on male sex determination gene in the chicken using genome editing technology
  3. 2016年12月01日, The 39th Annual Meeting of the Molecular Biology Society of Japan, Best Poster Award, The Molecular Biology Society of Japan, Are chicken epiblast-derived stem cells naïve or primed pluripotent state?

Patented

  1. Patent, EP:1264889, 2006/06/07
  2. Patent, US:7250275, 2007/07/31
  3. Patent, US:7619080, 2009/11/17
  4. Patent, US:7691589, 2010/04/06
  5. Patent, US:7771714, 2010/08/10
  6. Patent, US:8198082, 2012/06/12
  7. Patent, EP:2143791, 2014/04/23

Social Activities

Organizing Academic Conferences, etc.

  1. The 3rd Annual Meeting of The Japanese Society for Genome Editing, Organizaing committee member, 2017/07, 2018/07